An optional parameter is just tagged with an attribute. This attribute tells the compiler to insert the default value for that parameter at the call-site.
The call obj2.TestMethod();
is replaced by obj2.TestMethod(false);
when the C# code gets compiled to IL, and not at JIT-time.
So in a way it's always the caller providing the default value with optional parameters. This also has consequences on binary versioning: If you change the default value but don't recompile the calling code it will continue to use the old default value.
On the other hand, this disconnect means you can't always use the concrete class and the interface interchangeably.
You already can't do that if the interface method was implemented explicitly.