import os
## first file in current dir (with full path)
file = os.path.join(os.getcwd(), os.listdir(os.getcwd())[0])
file
os.path.dirname(file) ## directory of file
os.path.dirname(os.path.dirname(file)) ## directory of directory of file
...
And you can continue doing this as many times as necessary...
Edit: from os.path, you can use either os.path.split or os.path.basename:
dir = os.path.dirname(os.path.dirname(file)) ## dir of dir of file
## once you're at the directory level you want, with the desired directory as the final path node:
dirname1 = os.path.basename(dir)
dirname2 = os.path.split(dir)[1] ## if you look at the documentation, this is exactly what os.path.basename does.