[css] What does the "~" (tilde/squiggle/twiddle) CSS selector mean?

It is General sibling combinator and is explained in @Salaman's answer very well.

What I did miss is Adjacent sibling combinator which is + and is closely related to ~.

example would be

.a + .b {
  background-color: #ff0000;
}

<ul>
  <li class="a">1st</li>
  <li class="b">2nd</li>
  <li>3rd</li>
  <li class="b">4th</li>
  <li class="a">5th</li>
</ul>
  • Matches elements that are .b
  • Are adjacent to .a
  • After .a in HTML

In example above it will mark 2nd li but not 4th.

_x000D_
_x000D_
   .a + .b {_x000D_
     background-color: #ff0000;_x000D_
   }
_x000D_
<ul>_x000D_
  <li class="a">1st</li>_x000D_
  <li class="b">2nd</li>_x000D_
  <li>3rd</li>_x000D_
  <li class="b">4th</li>_x000D_
  <li class="a">5th</li>_x000D_
</ul>
_x000D_
_x000D_
_x000D_

JSFiddle